Affiliation entity
Équipe 2 PDM
Interdisplinary research theme
Anatomy, Zoology, Systematics, Phylogenetics, Evolutionary Biology, Paleontology, Morphology, Mammalogy
Evolutionary morphology of placental mammals, with a focus on South American taxa


guillaume.billet [at]
01 40 79 38 14
01 40 79 35 80
Custom address(es)

Bâtiment de paléontologie (CP38)

8 rue Buffon

75005 Paris


Responsibilities inside unit

Projet coordinator « Complex morphological variations and phylogenetic reconstructions »



My work aims at improving our understanding of the patterns of morphological evolution in deep time, using mostly the diversification of extinct and extant South American placentals as a model. My studies on these aspects follow three main objectives:

  • describing and analyzing the patterning of unusual morphological variations, at different hierarchical levels (eg, ontogenetical, interspecfic, inter-individual)

  • studying potential links between these patterns of morphological variation and functional characteristics

  • searching for a phylogenetic signal in morphological data and improving our use of this kind of data for phylogenetic reconstructions and placement of fossil taxa

Publication list


cv_billet.pdf Format (Pdf) - 244.84 KB


  • Mallet Christophe, Billet Guillaume, Cornette Raphaël & Alexandra Houssaye And, March 2022Adaptation to Graviportality in Rhinocerotoidea? An Investigation through the Long Bone Shape Variation in Their Hindlimb. , , dir. {Linnean Society of London}
  • Mallet Christophe, Houssaye Alexandra, Cornette Raphaël & Billet Guillaume, December 2021Long Bone Shape Variation in the Forelimb of Rhinocerotoidea: Relation with Size, Body Mass and Body Proportions. , , dir. {Linnean Society of London}
  • Boivin Myriam, MARIVAUX Laurent, Aguirre-Diaz Walter, Benites-Palomino Aldo, Billet Guillaume, Pujos François, Salas-Gismondi Rodolfo, Stutz Narla S, Tejada-Lara Julia V, Varas-Malca Rafael M, Walton Anne H & Antoine Pierre-Olivier, November 2021Late Middle Miocene Caviomorph Rodents from Tarapoto, Peruvian Amazonia. vol. 16, n° 11, dir. {Public Library of Science} e0258455
  • LE VERGER Kévin, González Ruiz Laureano R & Billet Guillaume, July 2021Comparative Anatomy and Phylogenetic Contribution of Intracranial Osseous Canals and Cavities in Armadillos and Glyptodonts (Xenarthra, Cingulata). , , dir. {Wiley}
  • Billet Guillaume & Bardin Jérémie, February 2021Segmental Series and Size: Clade-Wide Investigation of Molar Proportions Reveals a Major Evolutionary Allometry in the Dentition of Placental Mammals. , , dir. {Oxford University Press (OUP)}
  • Antoine Pierre-Olivier, Yans Johan, Castillo Angélica Aliaga, Stutz Narla, Abello M. Alejandra Alejandra, Adnet Sylvain, Custódio Michele Andriolli, Benites-Palomino Aldo, Billet Guillaume, Boivin Myriam, Herrera Fabiany, Jaramillo Carlos, Mártinez Camila, Moreno Federico, Navarrete Rosa E et al., 2021Biotic Community and Landscape Changes around the Eocene–Oligocene Transition at Shapaja, Peruvian Amazonia: Regional or Global Drivers?. , , dir. {Elsevier}
  • Marivaux Laurent, Aguirre-Diaz Walter, Benites-Palomino Aldo, Billet Guillaume, BOIVIN Myriam, Pujos Francois, Salas-Gismondi Rodolfo, Tejada-Lara Julia V., Varas-Malca Rafael & Antoine Pierre-Olivier, July 2020New Record of Neosaimiri (Cebidae, Platyrrhini) from the Late Middle Miocene of Peruvian Amazonia. vol. 146, , dir. {Elsevier} p. 102835
  • Mallet Christophe, Billet Guillaume, Houssaye Alexandra & Cornette Raphael, June 2020A First Glimpse at the Influence of Body Mass in the Morphological Integration of the Limb Long Bones: An Investigation in Modern Rhinoceroses. , , dir. {Wiley}
  • Vautrin Quentin, TABUCE Rodolphe, Lihoreau Fabrice, Bronnert Constance, Gheerbrant Emmanuel, Godinot Marc, Metais Gregoire, Yans Johan, Dutour Yves, Vialle Nicolas, Jean Philip & Billet Guillaume, 2020New Remains of Lophiaspis Maurettei (Mammalia, Perissodactyla) from the Early Eocene of France and the Implications for the Origin of the Lophiodontidae. vol. 40, n° 6, dir. {Society of Vertebrate Paleontology} e1878200
  • Gheerbrant Emmanuel, Schmitt Arnaud & Billet Guillaume, 2020Petrosal and Bony Labyrinth Morphology of the Stem Paenungulate Mammal (Paenungulatomorpha) Ocepeia Daouiensis from the Paleocene of Morocco. , , dir. {Wiley}